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1. 发明授权

US08664475B2 Plants with increased yield 失效
标题翻译：植物增产
公开(公告)号：US08664475B2
公开(公告)日：2014-03-04
申请号：US12678387
申请日：2008-09-17
申请人： Piotr Puzio , Oliver Bläsing , Oliver Thimm , Gerhard Ritte , Hardy Schön
发明人： Piotr Puzio , Oliver Bläsing , Oliver Thimm , Gerhard Ritte , Hardy Schön
IPC分类号： C12N15/82 , C12N15/63
CPC分类号： C12N15/8273 , C07K14/415 , C12N15/8261 , Y02A40/146
摘要： This invention relates generally to a plant cell with enhanced nitrogen use efficiency and/or increased biomass production as compared to a corresponding non-transformed wild type plant cell by increasing or generating one or more activities of polypeptides associated with enhanced nitrogen use efficiency in plants.
摘要翻译：本发明一般涉及与相应的未转化的野生型植物细胞相比，通过增加或产生与植物中提高的氮利用效率相关的多肽活性的一种或多种活性，具有增强的氮利用效率和/或增加的生物量产量的植物细胞。

2. 发明授权

US08809059B2 Plants with increased yield 有权
标题翻译：植物增产
公开(公告)号：US08809059B2
公开(公告)日：2014-08-19
申请号：US12678892
申请日：2008-09-19
申请人： Oliver Bläsing , Piotr Puzio , Oliver Thimm , Gerhard Ritte , Hardy Schön
发明人： Oliver Bläsing , Piotr Puzio , Oliver Thimm , Gerhard Ritte , Hardy Schön
IPC分类号： C12N15/82 , C12N5/04 , A01H5/00 , C12N15/87
CPC分类号： C12N15/8261 , C12N15/8271 , C12N15/8273 , Y02A40/146
摘要： The present invention disclosed herein provides a method for producing a plant with increased yield as compared to a corresponding wild type plant comprising increasing or generating one or more activities in a plant or a part thereof. The present invention further relates to nucleic acids enhancing or improving one or more traits of a transgenic plant, and cells, progenies, seeds and pollen derived from such plants or parts, as well as methods of making and methods of using such plant cell(s) or plant(s), progenies, seed(s) or pollen. Particularly, said improved trait(s) are manifested in an increased yield, preferably by improving one or more yield-related trait(s), e.g. low temperature tolerance.
摘要翻译：本文公开的本发明提供了与相应的野生型植物相比产生具有增加的产量的植物的方法，包括增加或产生植物或其部分中的一种或多种活性。本发明还涉及增强或改善一种或多种转基因植物，以及衍生自这种植物或部分的细胞，后代，种子和花粉的一种或多种性状的核酸，以及制备方法和使用这种植物细胞的方法）或植物，后代，种子或花粉。特别地，所述改良的性状表现为增加的产量，优选通过改善一种或多种产量相关性状，例如，耐低温。

3. 发明申请

US20110321197A1 Plants with Increased Yield (NUE) 审中-公开
标题翻译：提高产量的植物（NUE）
公开(公告)号：US20110321197A1
公开(公告)日：2011-12-29
申请号：US13125433
申请日：2009-10-02
申请人： Hardy Schön , Oliver Thimm , Gerhard Ritte , Oliver Bläsing , Koen Bruynseels , Yves Hatzfeld , Valerie Frankard , Ana Isabel Sanz Molinero , Christophe Reuzeau , Steven Vandenabeele
发明人： Hardy Schön , Oliver Thimm , Gerhard Ritte , Oliver Bläsing , Koen Bruynseels , Yves Hatzfeld , Valerie Frankard , Ana Isabel Sanz Molinero , Christophe Reuzeau , Steven Vandenabeele
IPC分类号： A01H1/06 , C12N15/63 , C12P21/00 , C12N9/88 , C12N9/12 , C12N9/10 , C12N9/00 , C12N9/04 , C12N9/16 , C12N9/22 , C12N9/90 , C07K2/00 , C07K16/40 , C07K16/14 , C07K16/12 , C12N5/10 , A01H5/00 , C12Q1/68 , G01N33/53 , C12Q1/02 , C12Q1/28 , C12Q1/32 , C12Q1/48 , C12Q1/44 , C12Q1/42 , C08B30/04 , A01C14/00 , C07H21/00
CPC分类号： C12N15/8273 , C12N15/8261 , Y02A40/146
摘要： A method for producing a plant with increased yield as compared to a corresponding wild type plant whereby the method comprises at least the following step: increasing or generating in a plant or a part thereof one or more activities selected from the group consisting of 17.6 kDa class I heat shock protein, 26.5 kDa class I small heat shock protein, 26S protease subunit, 2-Cys peroxiredoxin, 3-dehydroquinate synthase, 5-keto-D-gluconate-5-reductase, asparagine synthetase A, aspartate 1-decar-boxylase precursor, ATP-dependent RNA helicase, B0567-protein, B1088-protein, B1289-protein, B2940-protein, calnexin homolog, CDS5399-protein, chromatin structure-remodeling complex protein, D-amino acid dehydrogenase, D-arabinono-1,4-lac-tone oxidase, Delta 1-pyrroline-5-carboxylate reductase, glycine cleavage complex lipoylprotein, ketodeoxygluconokinase, lipoyl synthase, low-molecular-weight heat-shock protein, Microsomal cytochrome b reductase, mitochondrial ribosomal protein, mitotic check point protein, monodehydroascorbate reductase, paraquat-inducible protein B, phosphatase, Phosphoglucosamine mutase, protein disaggregation chaperone, protein kinase, pyruvate decarboxylase, recA family protein, rhodanese-related sulfurtransferase, ribonuclease P protein component, ribosome modulation factor, sensory histidine kinase, serine hydroxymethyltransferase, SLL1280-protein, SLL1797-protein, small membrane lipoprotein, Small nucleolar ribonucleoprotein complex subunit, Sulfatase, transcription initiation factor subunit, tretraspanin, tRNA ligase, xyloglucan galactosyltransferase, YKL130C-protein, YLR443W-protein, YML096W-protein, and zinc finger family protein-activity.
摘要翻译：与相应的野生型植物相比，生产具有增加的产量的植物的方法，其中所述方法至少包括以下步骤：在植物或其部分中增加或产生一种或多种选自17.6kDa类的活性 I热休克蛋白，26.5 kDa I类小热休克蛋白，26S蛋白酶亚基，2-Cys过氧化还原蛋白，3-脱氢秋水仙素合成酶，5-酮-D-葡萄糖酸-5-还原酶，天冬酰胺合成酶A，天冬氨酸1-脱羧基酶前体，ATP依赖性RNA解旋酶，B0567-蛋白，B1088-蛋白，B1289-蛋白，B2940蛋白，钙粘蛋白同源物，CDS5399-蛋白，染色质结构重塑复合蛋白，D-氨基酸脱氢酶，D-阿拉伯聚糖-1， 4-lac-色素氧化酶，1α-吡咯啉-5-羧酸还原酶，甘氨酸裂解复合物脂酰蛋白，酮脱氧葡萄糖激酶，脂酰基合成酶，低分子量热休克蛋白，微粒体细胞色素b还原酶，线粒体核糖体蛋白，有丝分裂检测点 nt蛋白，单脱氢抗坏血酸还原酶，百草枯诱导蛋白B，磷酸酶，磷酸葡糖胺变位酶，蛋白质分解伴侣蛋白，蛋白激酶，丙酮酸脱羧酶，recA家族蛋白，硫酸相关硫转移酶，核糖核酸酶P蛋白组分，核糖体调节因子，感觉组氨酸激酶，丝氨酸羟基甲基转移酶，SLL1280蛋白，SLL1797蛋白，小膜脂蛋白，小核仁核糖核蛋白复合子亚基，硫酸酯酶，转录起始因子亚基，tretraspanin，tRNA连接酶，木葡聚糖半乳糖基转移酶，YKL130C-蛋白，YLR443W蛋白，YML096W蛋白和锌指家庭蛋白质活性。

4. 发明申请

US20120227134A1 Plants with Increased Yield 审中-公开
标题翻译：植物增产
公开(公告)号：US20120227134A1
公开(公告)日：2012-09-06
申请号：US13510220
申请日：2010-11-05
申请人： Hardy Schön , Oliver Thimm , Gerhard Ritte , Oliver Bläsing , Stefan Henkes , Koen Bruynseels , Yves Hatzfeld , Valerie Frankard , Ana Isabel Sanz Molinero , Christophe Reuzeau , Steven Vandenabeele , Bryan McKersie , Krishna Kolliparra , Christian Dammann
发明人： Hardy Schön , Oliver Thimm , Gerhard Ritte , Oliver Bläsing , Stefan Henkes , Koen Bruynseels , Yves Hatzfeld , Valerie Frankard , Ana Isabel Sanz Molinero , Christophe Reuzeau , Steven Vandenabeele , Bryan McKersie , Krishna Kolliparra , Christian Dammann
IPC分类号： A01H5/00 , C12P21/00 , C12N5/10 , C12Q1/48 , C12Q1/42 , C12Q1/26 , A01H1/06 , C12Q1/68 , C12N15/52 , C12N15/29 , C07K14/415 , C07K16/16 , A01H5/10 , G01N33/566 , A01N25/00 , A01N43/00 , A01N37/00 , C12N15/82
CPC分类号： C12N15/8261 , C07K14/415 , Y02A40/146
摘要： A method for producing a plant with increased yield as compared to a corresponding wild type plant whereby the method comprises at least the following step: increasing or generating in a plant or a part thereof one or more activities of a polypeptide selected from the group consisting of 2-oxoglutarate-dependent dioxygenase, 3-ketoacyl-CoA thiolase, 3′-phosphoadenosine 5′-phosphate phosphatase, 4-diphosphocytidyl-2-C-methyl-D-erythritol kinase, 5OS chloroplast ribosomal protein L21, 57972199. R01.1-protein, 60952769. R01.1-protein, 60S ribosomal protein, ABC transporter family protein, AP2 domain-containing transcription factor, argonaute protein, AT1 G29250.1-protein, AT1 G53885-protein, AT2G35300-protein, AT3G04620-protein, AT4G01870-protein, AT5G42380-protein, AT5G47440-protein, CDS5394-protein, CDS5401_TRUNCATED-protein, cold response protein, cullin, Cytochrome P450, delta-8 sphingolipid desaturase, galactinol synthase, glutathione-S-transferase, GTPase, haspin-related protein, heat shock protein, heat shock transcription factor, histone H2B, jasmonate-zim-domain protein, mitochondrial asparaginyl-tRNA synthetase, Oligosaccharyltransferase, OS02G44730-protein, Oxygen-evolving enhancer protein, peptidyl-prolyl cis-trans isomerase, peptidyl-prolyl cis-trans isomerase family protein, plastid lipid-associated protein, Polypyrimidine tract binding protein, PRLI-interacting factor, protein kinase, protein kinase family protein, rubisco subunit binding-protein beta subunit, serine acetyltransferase, serine hydroxymethyltransferase, small heat shock protein, S-ribosylhomocysteinase, sugar transporter, Thioredoxin H-type, ubiquitin-conjugating enzyme, ubiquitin-protein ligase, universal stress protein family protein, and Vacuolar protein.
摘要翻译：与相应的野生型植物相比，生产具有增加的产量的植物的方法，其中所述方法至少包括以下步骤：在植物或其部分中增加或产生一种或多种选自以下的多肽的活性： 2-氧戊二酸依赖性双加氧酶，3-酮酰基辅酶A硫解酶，3'-磷酸腺苷5'-磷酸磷酸酶，4-二磷酸胞苷-2-C-甲基-D-赤藓糖醇激酶，5OS叶绿体核糖体蛋白L21,57972199。R01.1 蛋白质，60952769.R01.1-蛋白质，60S核糖体蛋白质，ABC转运蛋白家族蛋白质，含AP2结构域的转录因子，argonaute蛋白质，AT1 G29250.1-蛋白质，AT1 G53885-蛋白质，AT2G35300-蛋白质，AT3G04620-蛋白质， AT4G01870蛋白质，AT5G42380蛋白质，AT5G47440蛋白质，CDS5394蛋白质，CDS5401_TRUNCATED蛋白质，冷应答蛋白质，cullin，细胞色素P450，δ-8鞘脂类去饱和酶，肌动蛋白醇合酶，谷胱甘肽-S-转移酶，GTP酶，，热休克蛋白，热休克转录因子，组蛋白H2B，茉莉酸-zim结构域蛋白，线粒体天冬酰胺酰-tRNA合成酶，寡糖转移酶，OS02G44730-蛋白，氧进化增强子蛋白，肽基脯氨酰顺反异构酶，肽基脯氨酰顺式 - 反式异构酶家族蛋白质，质体脂质相关蛋白，聚嘧啶多肽结合蛋白，PRLI相互作用因子，蛋白激酶，蛋白激酶家族蛋白，rubisco亚基结合蛋白β亚基，丝氨酸乙酰转移酶，丝氨酸羟甲基转移酶，小热休克蛋白，S - 糖基半胱氨酸酶，糖转运蛋白，硫氧还蛋白H型，泛素结合酶，泛素 - 蛋白连接酶，通用应激蛋白家族蛋白和Vacuolar蛋白。

5. 发明申请

US20110195843A1 Plants with Increased Yield (LT) 审中-公开
标题翻译：提高产量的植物（LT）
公开(公告)号：US20110195843A1
公开(公告)日：2011-08-11
申请号：US13120279
申请日：2009-09-18
申请人： Oliver Thimm , Gerhard Ritte , Hardy Schön , Oliver Bläsing
发明人： Oliver Thimm , Gerhard Ritte , Hardy Schön , Oliver Bläsing
IPC分类号： A01N61/00 , A01H1/00 , C07H21/00 , C12N15/82 , C12P21/00 , C07K2/00 , C07K16/00 , C12N1/00 , C12N5/10 , A01H5/00 , G01N33/53 , A01P21/00 , A01G1/00 , A01C14/00
CPC分类号： C12N15/8261 , C07K14/245 , Y02A40/146
摘要： A method for producing a plant with increased yield as compared to a corresponding wild type plant comprising at least the following step: increasing or generating in a plant or a part thereof one or more activities selected from the group consisting of 3-phosphoglycerate dehydrogenase, Adenylate kinase, B2758-protein, Cyclic nucleotide phosphodiesterase, cysteine synthase, Exopolyphosphatase, geranylgeranyl reductase, Mating hormone A-factor precursor, mitochondrial succinate-fumarate transporter, modification methylase HemK family protein, Myo-inositol transporter, oxidoreductase subunit, peptidy-prolyl-cis-trans-isomerase, protein kinase, Ribose-5-phosphate isomerase, slr1293-protein, YDR049W-protein, YJL181W-protein, and YPL109C-protein-activity.
摘要翻译：与相应的野生型植物相比，产生具有增加的产量的植物的方法，其包括至少以下步骤：在植物或其部分中增加或产生一种或多种选自3-磷酸甘油酸脱氢酶，腺苷酸激酶，B2758-蛋白，环核苷酸磷酸二酯酶，半胱氨酸合成酶，外多糖磷酸酶，香叶基香叶基还原酶，交配激素A因子前体，线粒体琥珀酸 - 富马酸转运蛋白，修饰甲基化酶HemK家族蛋白，肌醇转运蛋白，氧化还原酶亚基，肽 - 脯氨酰 - 顺式蛋白激酶，核糖-5-磷酸异构酶，slr1293蛋白，YDR049W蛋白，YJL181W蛋白和YPL109C蛋白活性。

6. 发明申请

US20110010800A1 PLANTS WITH INCREASED YIELD 审中-公开
标题翻译：植物增加
公开(公告)号：US20110010800A1
公开(公告)日：2011-01-13
申请号：US12919507
申请日：2009-02-27
申请人： Gerhard Ritte , Oliver Bläsing , Oliver Thimm
发明人： Gerhard Ritte , Oliver Bläsing , Oliver Thimm
IPC分类号： A01H1/06 , C12Q1/68 , C12Q1/02 , C12P21/02 , C12N5/10 , C12N15/82 , A01N43/00 , C07K14/245 , C07K16/00 , C07H21/00 , A01H5/00 , A01H5/10 , C07K14/39
CPC分类号： C12N15/8261 , C07K14/415 , Y02A40/146
摘要： The present invention disclosed herein provides a method for producing a plant with increased yield as compared to a corresponding wild type plant comprising increasing or generating one or more activities in a plant or a part thereof. The present invention further relates to nucleic acids enhancing or improving one or more traits of a transgenic plant, and cells, progenies, seeds and pollen derived from such plants or parts, as well as methods of making and methods of using such plant cell(s) or plant(s), progenies, seed(s) or pollen. Particularly, said improved trait(s) are manifested in an increased yield, preferably by improving one or more yield-related trait(s).
摘要翻译：本文公开的本发明提供了与相应的野生型植物相比产生具有增加的产量的植物的方法，包括增加或产生植物或其部分中的一种或多种活性。本发明还涉及增强或改善一种或多种转基因植物，以及衍生自这种植物或部分的细胞，后代，种子和花粉的一种或多种性状的核酸，以及制备方法和使用这种植物细胞的方法）或植物，后代，种子或花粉。特别地，所述改良的性状表现为增加的产量，优选通过改善一种或多种产量相关性状。

7. 发明申请

US20110277179A1 Plants With Increased Yield 失效
标题翻译：植物增产
公开(公告)号：US20110277179A1
公开(公告)日：2011-11-10
申请号：US12678387
申请日：2008-09-17
申请人： Piotr Puzio , Oliver Bl¿sing , Oliver Thimm , Gerhard Ritte , Hardy Sch¿n
发明人： Piotr Puzio , Oliver Bl¿sing , Oliver Thimm , Gerhard Ritte , Hardy Sch¿n
IPC分类号： A01H5/00 , A01H1/00 , C12N5/10 , C12N15/29 , C12Q1/68 , C12N15/82
CPC分类号： C12N15/8273 , C07K14/415 , C12N15/8261 , Y02A40/146
摘要： This invention relates generally to a plant cell with enhanced nitrogen use efficiency and/or increased biomass production as compared to a corresponding non-transformed wild type plant cell by increasing or generating one or more activities of polypeptides associated with enhanced nitrogen use efficiency in plants.
摘要翻译：本发明一般涉及与相应的未转化的野生型植物细胞相比，通过增加或产生与植物中提高的氮利用效率相关的多肽活性的一种或多种活性，具有增强的氮利用效率和/或增加的生物量产量的植物细胞。

8. 发明申请

US20120117867A1 Plants with Increased Yield 审中-公开
标题翻译：植物增产
公开(公告)号：US20120117867A1
公开(公告)日：2012-05-17
申请号：US13386273
申请日：2010-07-15
申请人： Janneke Hendriks , Oliver Thimm , Philip Groth , Alexandre Prokoudine , Gerhard Ritte , Claudia König , Resham Kulkarni , Krishna Kollipara
发明人： Janneke Hendriks , Oliver Thimm , Philip Groth , Alexandre Prokoudine , Gerhard Ritte , Claudia König , Resham Kulkarni , Krishna Kollipara
IPC分类号： A01G1/00 , C12Q1/68 , C12Q1/02 , C12P21/06 , C12N5/10 , C12N15/82 , A01N43/54 , A01N37/00 , C07K14/00 , C07K16/00 , C07H21/04 , C08B31/00 , A01H5/00 , C07K14/245 , C07K14/39 , A01H5/10 , A01C14/00 , C10L1/00
CPC分类号： C12N15/8261 , C12N15/8271 , C12N15/8273 , Y02A40/146
摘要： A method for producing a plant with increased yield as compared to a corresponding wild type plant whereby the method comprises at least the following step: increasing or generating in a plant or a part thereof one or more activities of a polypeptide selected from the group consisting of 26S proteasome-subunit, 50S ribosomal protein L36, Autophagy-related protein, B0050-protein, Branched-chain amino acid permease, Calmodulin, carbon storage regulator, FK506-binding protein, gamma-glutamyl-gamma-aminobutyrate hydrolase, GM02LC38418-protein, Heat stress transcription factor, Mannan polymerase II complex subunit, mitochondrial precursor of Lon protease homolog, MutS protein homolog, phosphate transporter subunit, Protein EFR3, pyruvate kinase, tellurite resistance protein, Xanthine permease, and YAR047C-protein.
摘要翻译：与相应的野生型植物相比，生产具有增加的产量的植物的方法，其中所述方法至少包括以下步骤：在植物或其部分中增加或产生一种或多种选自以下的多肽的活性： 26S蛋白酶体亚基，50S核糖体蛋白L36，自噬相关蛋白，B0050蛋白，支链氨基酸通透酶，钙调蛋白，碳储存调节剂，FK506结合蛋白，γ-谷氨酰-γ-氨基丁酸酯水解酶，GM02LC38418蛋白，热应激转录因子，Mannan聚合酶II复合亚单位，Lon蛋白酶同源物的线粒体前体，MutS蛋白同源物，磷酸转运子亚基，蛋白EFR3，丙酮酸激酶，碲化物抗性蛋白，黄嘌呤通透酶和YAR047C蛋白。

9. 发明授权

US08269063B2 Methods for identifying proteins with starch phosphorylating enzymatic activity 有权
标题翻译：用淀粉磷酸化酶活性鉴定蛋白质的方法
公开(公告)号：US08269063B2
公开(公告)日：2012-09-18
申请号：US10591419
申请日：2005-03-04
申请人： Claus Frohberg , Oliver Koetting , Gerhard Ritte , Martin Steup
发明人： Claus Frohberg , Oliver Koetting , Gerhard Ritte , Martin Steup
IPC分类号： C12N15/82 , C12N15/00 , C12N15/10 , C07H21/04
CPC分类号： C12N9/12 , C12N15/8245 , C12Q1/485 , G01N33/573 , G01N2400/14 , G01N2500/04
摘要： The present invention relates to a method for identifying proteins involved in the phosphorylation of starch and nucleic acids which code for such proteins. The present invention further relates to plant cells and plants which exhibit an altered activity of a protein which can be identified using the method according to the invention. Plant cells and plants of this type synthesise a modified starch. The present invention therefore also relates to the starch synthesised by the plant cells and plants according to the invention as well as to methods for the manufacture of this starch and to the manufacture of starch derivatives of this modified starch.
摘要翻译：本发明涉及一种鉴定涉及淀粉磷酸化的蛋白质的方法以及编码这种蛋白质的核酸。本发明还涉及使用本发明方法鉴定的蛋白质活性变化的植物细胞和植物。这种植物细胞和植物合成改性淀粉。因此，本发明还涉及由根据本发明的植物细胞和植物合成的淀粉以及制备该淀粉的方法和该淀粉衍生物的制造。

10. 发明授权

US07772463B2 Plants with increased activity of a starch phosphorylating enzyme 有权
标题翻译：具有增加淀粉磷酸化酶活性的植物
公开(公告)号：US07772463B2
公开(公告)日：2010-08-10
申请号：US10591428
申请日：2005-03-04
申请人： Claus Frohberg , Oliver Koetting , Gerhard Ritte , Martin Steup
发明人： Claus Frohberg , Oliver Koetting , Gerhard Ritte , Martin Steup
IPC分类号： C12N15/82 , C12N15/00 , C12N15/29 , C12N15/52 , C12N15/63 , A01H5/00
CPC分类号： B02C9/00 , C12N9/12 , C12N9/1205 , C12N15/8245
摘要： The present invention relates to plant cells and plants, which are genetically modified, whereby the genetic modification leads to an increase in the activity of a starch-phosphorylating OK1 protein in comparison to the corresponding wild type plant cells or wild type plants that have not been genetically modified. In addition, the present invention concerns means and methods for the manufacture of such plant cells and plants. These types of plant cells and plants synthesize a modified starch. Therefore, the present invention also concerns the starches synthesized from the plant cells and plants according to the invention, methods for manufacturing these starches, and the manufacture of starch derivatives of these modified starches, as well as flours containing starches according to the invention.Furthermore, the present invention also relates to nucleic acids, coding starch-phosphorylating OK1 proteins, vectors, host cells, plant cells, and plants containing such nucleic acid molecules. In addition, the present invention relates to OK1 proteins that have starch-phosphorylating activity.
摘要翻译：本发明涉及遗传修饰的植物细胞和植物，其中遗传修饰导致淀粉磷酸化OK1蛋白的活性与尚未被使用的相应野生型植物细胞或野生型植物相比增加转基因此外，本发明涉及用于制造这种植物细胞和植物的方法和方法。这些类型的植物细胞和植物合成改性淀粉。因此，本发明还涉及由根据本发明的植物细胞和植物合成的淀粉，这些淀粉的制造方法以及这些改性淀粉的淀粉衍生物的制备，以及含有本发明淀粉的面粉。此外，本发明还涉及编码淀粉磷酸化OK1蛋白，载体，宿主细胞，植物细胞和含有该核酸分子的植物的核酸。此外，本发明涉及具有淀粉磷酸化活性的OK1蛋白质。

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